Table 1 Notation General remarks

A

modifying locus with two alleles A ₁ and A ₂

B

object locus with two alleles B ₁ and B ₂

two-locus gamete (A _i B _k ) with allele i at the A- and allele k at the

B-locus

zygote or any other diploid genotype originating from fusion of ga-

metes (A _i B _k ) and (A _j B _l ). Note that because of unordered geno-

types but because of linkage

a _i , b _k

relative frequencies of alleles A _i , B _k among adults

relative frequency of gamete in the gametic production

relative frequency of genotype among adults

a _ij , b _kl

relative frequency of genotype A _i A _j and B _k B _l , respectively, among

adults

r

recombination frequency (0 ≤ r ≤ ½)

probability of fusion when the gametes and encounter.

f-values are assumed to differ in only three ways:

f ₂ := probability of fusion in the presence of allele A ₂ in at least

one of the two encountering gametes, for all i, k, l;

:= probability of fusion when A ₁-carriers with equal B-alleles

encounter (homotypic mating), ;

:= fusion probability among A ₁-carriers with different B-

alleles (heterotypic mating), ;

^v kl

viabilities at the B-locus

combined selection value of fusion probability and viability selection,

defined as